LRP5 FIGF ACTR1A integrin OMD TIEG TGFb signaling DLX2 TGFB1I4 EP300 ZFHX1B PKA PRKACB BMP4 FGF2 adipocytes PPARG ZBTB17 MIZ1 KLF3 PRKAR2A NAB2 NAB1 RBL1 NCOR2 NCOA3 JUND NR4A1 TWIST1 RUNX2 ATF1 actin MAPK3 CREBL1 BARX1 DLX5 Osterix HIPK2 RARG CCNB2 CCNA2 CDK2 BMP2 DCN COL6A1 BGN FGF7 TIMP2 catenin COL5A1 COL5A3